Classification, reconstructed phylogeny and geographical history of the New World species of Coptodera Dejean (Coleoptera: Carabidae: Lebiini)
Proceedings of the Entomological Society of Ontario
1993
Journal Article
124
1-182
Parque Nacional del Manu morphology systematics Coleoptera Insecta Arthropoda Invertebrata Animalia new species taxonomy Coptodera Pakitza animals arthropods insects invertebrates beetles Madre de Dios Bibliography
Keys, descriptions, and illustrations distinguish the known New World species of Coptodera (type species-Coptodera festiva Dejean), and each taxon is characterized in terms of structural features of adults, habitat, geographical distribution, chorological affinities, and phylogenetic relationships. Twelve new species are described. The New World species of Coptodera are arranged in eleven groups, sequenced according to the reconstructed phylogeny: C. sallei group, including C. sallei new name; C. elongata group, including C. elongata Putzeys, C. schaumi Chaudoir, C. megalops Bates, C. championi Bates, and C. apicalis new species (type locality-Ecuador, Esmeraldas, Zapallo Grande); C. aurata group, including C. xanthopleura Bates, C. aurata Chevrolat, and C. viridis new species (type locality-Mexico, Colima, near Ixtapa); C. depressa group" including C. depressa Dejean and C. versicolor Bates; C. transversa group, including C. dromioides (Bates), C. nigrosignata (Chaudoir); C. fulminans (Bates), C. nigrostriata (Reiche), C. lineata (Bates), C. stockwelli new species (type locality-Panama, Canal Zone, Madden Forest), C. transversa (Reiche), C. sigillata new species (type locality-Brazil, Santa Catarina, Nova Teutonia), and C. squiresi (Chaudoir); C. festiva group, including C. sahlbergi Chaudoir, C. relucens Bates, C. poecila Bates, C. pakitza new species (type locality-Peru, Madre de Dios, Pakitza), C. festiva Dejean, C. foveolata new species (type locality-Mexico, Veracruz, Tebanca), C. nigroviridis new species (type locality-Mexico, Chiapas, Parque Montebello), C. rufescens Buquet, and C. teutonica new species (type locality-Brazil, Santa Catarina, Nova Teutonia); C. aeneorufa group, including C. aeneorufa Bates; C. bifasciata group, including C. bifasciata Putzeys, C. tripartita Chaudoir, C. cupreotincta Bates, and C. braziliensis new species (type locality-Brazil, Santa Catarina, Hansa Humboldt); C. emarginata group, including C. chalcites Bates, C. emarginata Dejean, and C. acutipennis (Buquet); C. picea group, including C. picea Dejean; and C. aerata group, including C. nitidula (Buquet), C. brunnea new species (type locality-United States, Arizona, Cochise County, Guadalupe Canyon), C. aerata Dejean, C. waytkowskii Liebkc, C. erwini new species (type locality-Peru, Madre de Dios, Pakitza), and C. tripunctata new species (type locality-Peru, Loreto, Cocha Shinguito). Removed from synonymy with C. lineata Chaudoir is C. nigrosignata Chaudoir, the name of a valid species. Ten synonymies are newly proposed with the senior synonym of each combination listed first Coptodera Dejean = Stenoglossa Chaudoir, C. megalops Bates = C. pertzeli Liebke = C. peruana Liebke; C. depressa Dejean = C. plaumanni Liebke; C. sahlbergi Chaudoir = C. discicollis Chaudoir, C. festiva Dejean C. chloris Bates; C. cupreotincta Bates = C. tesselatus Chaudoir; C. acutipennis (Buquet) = C. scintillans Bates; C. picea Dejean = C. unicolor Chevrolat; C. nitidula (Buquet) = C. triangularis (Buquet) = C. nubiculosa Chaudoir = C. flavodisca Chaudoir; C. lineata (Bates) = C. nigrosignata (Chaudoir); and C. transversa (Reiche) = C. picturata (Chaudoir) = C. undulata (Chaudoir). One replacement name is proposed for a junior homonym: C. sallei for C. variegata Chaudoir. Six species are removed from Coptodera and placed into other genera in the following new combinations: Microlestodes marcida Blackburn, Anchista philippinensis Jedlicka, Stenognathus robustus Bates, Lelis polygona Bates, Lelis latipennis Bates, and Lelis rutila Bates. Coptodera fasciolata Macleay is given new status as the previously used name, C. aurata Macleay, and synonym, C. tesselatus Chaudoir (1883), are both junior homonyms of C. aurata Chevrolat and C. tesselatus Chaudoir (1869) respectively.